The presenters take on a tour of Northern Italy, travelling across the country in a car of their choosing — May takes the Rolls-Royce Dawn, Clarkson takes the Aston Martin DB11, and Hammond takes the Dodge Challenger SRT Hellcat. The group begin at the Palio di Siena horse race, exploring art in Florence, racing each other at the Mugello Circuit, visiting the Lamborghini Museum and Piazza dei Signori, Vicenza, and finish off their tour in the city of Venice. Meanwhile Hammond and May fulfil a bet from the first episode during their hypercar reviews, by destroying Clarkson's old home in Chadlington, Oxfordshire, as part of the wager their colleague lost.

The presenters head to Switzerland to compare a supercar of the past, present and future to see which is best — Clarkson represents the past with a petrol powered Lamborghini Aventador S, May represents the present with a hybrid powered Honda NSX, and Hammond represents the future with an all-electric powered Rimac Concept One — taking them on a journey across the Swiss Alps, where they check out museums, conduct a drag race, and then put their cars into a hill climbing race at Hemberg. In a new segment entitled "Celebrity Face Off", Ricky Wilson and David Hasselhoff perform a timed lap each to see who is the fastest between them.

In the second part of their Colombia road trip, the trio make modifications to their vehicles in Bogotá and partake in a football match with their Colombian fixers during the time. As they head southwards for the remaining animals on their list, they find themselves contending with mountains, higher altitudes and severe weather, before their search for the final animal is found to be much easier than expected

The presenters head to Nevada for a RV holiday, buying their own and modifying it for a journey across roads and desert terrain, testing them in a series of challenging conditions before racing them in a demolition derby. May converts a National RV Tropi-Cal 6350 into a fully functional English pub, Clarkson transforms a Fleetwood Pace Arrow to appear more yacht-like, complete with a flying bridge on the roof, and Hammond chooses a minimalist approach by installing a tent on an International Harvester flatbed truck. Along the way, the group have a go at dune buggy racing, and conduct a comparison test with three American cars at the Spring Mountain race complex — the Chevrolet Corvette ZR-1 tested by Clarkson, the Jeep Grand Cherokee Trackhawk tested by Hammond and the Cadillac CTS-V tested by May.

The presenters pay tribute to the mid-size Ford saloon, with Clarkson and May looking over the history of the Ford Cortina and the generation of cars modeled on the original, while Hammond reviews the Ford Sierra RS Cosworth at the Eboladrome, before joining his colleagues to bid farewell to the Ford Mondeo with a car meet at Lincoln Cathedral. In their final scene, the group pay homage to their involvement in studio based car shows over the last 17 years, since the reboot of Top Gear.

The trio travel through Norway, Sweden and Finland in three Rally-Inspired Sports Saloons—Hammond purchases a 2004 Subaru Impreza WRX STI, May purchases a 2004 Mitsubishi Lancer Evolution VIII, and Clarkson purchases a 2007 Audi RS4. As they venture across Scandinavia towards the Russian border, they put their cars through their paces in a series of harsh winter tests.