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[3] Confirmed H. habilis fossils are dated to between 2.1 and 1.5 million years ago. This date range overlaps with the emergence of Homo erectus.^[1]^[2]

[4] Hominins with "proto-Homo" traits may have lived as early as 2.8 million years ago, as suggested by a fossil jawbone classified as transitional between Australopithecus and Homo discovered in 2015.

[6] A species proposed in 2010 based on the fossil remains of three individuals dated between 1.9 and 0.6 million years ago. The same fossils were also classified as H. habilis, H. ergaster or Australopithecus by other anthropologists.

[8] H. erectus may have appeared some 2 million years ago. Fossils dated to as much as 1.8 million years ago have been found both in Africa and in Southeast Asia, and the oldest fossils by a narrow margin (1.85 to 1.77 million years ago) were found in the Caucasus, so that it is unclear whether H. erectus emerged in Africa and migrated to Eurasia, or if, conversely, it evolved in Eurasia and migrated back to Africa.

[11] Homo erectus soloensis, found in Java, is considered the latest known survival of H. erectus. Formerly dated to as late as 50,000 to 40,000 years ago, a 2011 study pushed back the date of its extinction of H. e. soloensis to 143,000 years ago at the latest, more likely before 550,000 years ago. ^[6]

[12] Now also included in H. erectus are Peking Man (formerly Sinanthropus pekinensis) and Java Man (formerly Pithecanthropus erectus).

[13] H. erectus is now grouped into various subspecies, including Homo erectus erectus, Homo erectus yuanmouensis, Homo erectus lantianensis, Homo erectus nankinensis, Homo erectus pekinensis, Homo erectus palaeojavanicus, Homo erectus soloensis, Homo erectus tautavelensis, Homo erectus georgicus. The distinction from descendant species such as Homo ergaster, Homo floresiensis, Homo antecessor, Homo heidelbergensis and indeed Homo sapiens is not entirely clear.

[15] The type fossil is Mauer 1, dated to ca. 0.6 million years ago. The transition from H. heidelbergensis to H. neanderthalensis between 300 and 243 thousand years ago is conventional, and makes use of the fact that there is no known fossil in this period. Examples of H. heidelbergensis are fossils found at Bilzingsleben (also classified as Homo erectus bilzingslebensis).

[22] The age of H. sapiens has long been assumed to be close to 200,000 years, but since 2017 there have been a number of suggestions extending this time to as high as 300,000 years. In 2017, fossils found in Jebel Irhoud (Morocco) suggest that Homo sapiens may have speciated by as early as 315,000 years ago.^[12] Genetic evidence has been adduced for an age of roughly 270,000 years.^[13]

[25] The first humans with "proto-Neanderthal traits" lived in Eurasia as early as 0.6 to 0.35 million years ago (classified as H. heidelbergensis, also called a chronospecies because it represents a chronological grouping rather than being based on clear morphological distinctions from either H. erectus or H. neanderthalensis). There is a fossil gap in Europe between 300 and 243 kya, and by convention, fossils younger than 243 kya are called "Neanderthal".^[15]

[27] younger than 450 kya, either between 190–130 or between 70–10 kya^[16]

[30] rovisional names Homo sp. Altai or Homo sapiens ssp. Denisova.

[1] [1]
Schrenk F, Kullmer O, Bromage T (2007). "The Earliest Putative Homo Fossils". In Henke W, Tattersall I (eds.). Handbook of Paleoanthropology. Vol. 1. In collaboration with Thorolf Hardt. Berlin, Heidelberg: Springer. pp. 1611–1631. doi:10.1007/978-3-540-33761-4_52. ISBN 978-3-540-32474-4.

[2] [2]
DiMaggio EN, Campisano CJ, Rowan J, Dupont-Nivet G, Deino AL, Bibi F, et al. (March 2015). "Paleoanthropology. Late Pliocene fossiliferous sedimentary record and the environmental context of early Homo from Afar, Ethiopia". Science. 347 (6228): 1355–9. Bibcode:2015Sci...347.1355D. doi:10.1126/science.aaa1415. PMID 25739409. S2CID 43455561.

[5] [3]
Curnoe D (June 2010). "A review of early Homo in southern Africa focusing on cranial, mandibular and dental remains, with the description of a new species (Homo gautengensis sp. nov.)". Homo. 61 (3): 151–77. doi:10.1016/j.jchb.2010.04.002. PMID 20466364.

[7] [4]
Haviland WA, Walrath D, Prins HE, McBride B (2007). Evolution and Prehistory: The Human Challenge (8th ed.). Belmont, CA: Thomson Wadsworth. p. 162. ISBN 978-0-495-38190-7.

[9] [5]
Ferring R, Oms O, Agustí J, Berna F, Nioradze M, Shelia T, et al. (June 2011). "Earliest human occupations at Dmanisi (Georgian Caucasus) dated to 1.85-1.78 Ma". Proceedings of the National Academy of Sciences of the United States of America. 108 (26): 10432–6. Bibcode:2011PNAS..10810432F. doi:10.1073/pnas.1106638108. PMC 3127884. PMID 21646521.

[10] [6]
Indriati E, Swisher CC, Lepre C, Quinn RL, Suriyanto RA, Hascaryo AT, et al. (2011). "The age of the 20 meter Solo River terrace, Java, Indonesia and the survival of Homo erectus in Asia". PLOS ONE. 6 (6): e21562. Bibcode:2011PLoSO...621562I. doi:10.1371/journal.pone.0021562. PMC 3126814. PMID 21738710.

[Hazarika2007-14] [7]
Hazarika M (2007). "Homo erectus/ergaster and Out of Africa: Recent Developments in Paleoanthropology and Prehistoric Archaeology" (PDF). EAA Summer School eBook. Vol. 1. European Anthropological Association. pp. 35–41. Intensive Course in Biological Anthrpology, 1st Summer School of the European Anthropological Association, 16–30 June, 2007, Prague, Czech Republic

[16] [8]
Muttoni G, Scardia G, Kent DV, Swisher CC, Manzi G (2009). "Pleistocene magnetochronology of early hominin sites at Ceprano and Fontana Ranuccio, Italy". Earth and Planetary Science Letters. 286 (1–2): 255–268. Bibcode:2009E&PSL.286..255M. doi:10.1016/j.epsl.2009.06.032.

[17] [9]
Ji Q, Wu W, Ji Y, Li Q, Ni X (25 June 2021). "Late Middle Pleistocene Harbin cranium represents a new Homo species". The Innovation. 2 (3): 100132. doi:10.1016/j.xinn.2021.100132. PMC 8454552. PMID 34557772.

[18] [10]
Ni X, Ji Q, Wu W, Shao Q, Ji Y, Zhang C, Liang L, Ge J, Guo Z, Li J, Li Q, Grün R, Stringer C (25 June 2021). "Massive cranium from Harbin in northeastern China establishes a new Middle Pleistocene human lineage". The Innovation. 2 (3): 100130. doi:10.1016/j.xinn.2021.100130. PMC 8454562. PMID 34557770.

[eLIFE-2017a-19] [11]
Dirks PH, Roberts EM, Hilbert-Wolf H, Kramers JD, Hawks J, Dosseto A, et al. (May 2017). "Homo naledi and associated sediments in the Rising Star Cave, South Africa". eLife. 6: e24231. doi:10.7554/eLife.24231. PMC 5423772. PMID 28483040.

[20] [12]
Callaway, Ewan (7 June 2017). "Oldest Homo sapiens fossil claim rewrites our species' history". Nature. doi:10.1038/nature.2017.22114. Retrieved 11 June 2017.

[21] [13]
Posth C, Wißing C, Kitagawa K, Pagani L, van Holstein L, Racimo F, et al. (July 2017). "Deeply divergent archaic mitochondrial genome provides lower time boundary for African gene flow into Neanderthals". Nature Communications. 8: 16046. Bibcode:2017NatCo...816046P. doi:10.1038/ncomms16046. PMC 5500885. PMID 28675384.

[23] [14]
Bischoff JL, Shamp DD, Aramburu A, et al. (March 2003). "The Sima de los Huesos Hominids Date to Beyond U/Th Equilibrium (>350 kyr) and Perhaps to 400–500 kyr: New Radiometric Dates". Journal of Archaeological Science. 30 (3): 275–280. doi:10.1006/jasc.2002.0834. ISSN 0305-4403.

[24] [15]
Dean D, Hublin JJ, Holloway R, Ziegler R (May 1998). "On the phylogenetic position of the pre-Neandertal specimen from Reilingen, Germany". Journal of Human Evolution. 34 (5): 485–508. doi:10.1006/jhev.1998.0214. PMID 9614635.

[Changetal2015-26] [16]
Chang CH, Kaifu Y, Takai M, Kono RT, Grün R, Matsu'ura S, et al. (January 2015). "The first archaic Homo from Taiwan". Nature Communications. 6: 6037. Bibcode:2015NatCo...6.6037C. doi:10.1038/ncomms7037. PMC 4316746. PMID 25625212.

[Détroit2019-28] [17]
Détroit F, Mijares AS, Corny J, Daver G, Zanolli C, Dizon E, et al. (April 2019). "A new species of Homo from the Late Pleistocene of the Philippines" (PDF). Nature. 568 (7751): 181–186. Bibcode:2019Natur.568..181D. doi:10.1038/s41586-019-1067-9. PMID 30971845. S2CID 106411053.

[NYT-20190410-29] [18]
Zimmer C (10 April 2019). "A new human species once lived in this Philippine cave – Archaeologists in Luzon Island have turned up the bones of a distantly related species, Homo luzonensis, further expanding the human family tree". The New York Times. Retrieved 10 April 2019.

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Lineages	Temporal range (kya)	Habitat	Adult height	Adult mass	Cranial capacity (cm³)	Fossil record	Discovery	Publication of name
H. habilis membership in Homo uncertain	2,100–1,500^{[lower-alpha 1]}^{[lower-alpha 2]}	Tanzania	110–140 cm (3 ft 7 in – 4 ft 7 in)	33–55 kg (73–121 lb)	510–660	Many	1960	1964
H. rudolfensis membership in Homo uncertain	1,900	Kenya			700	2 sites	1972	1986
H. gautengensis also classified as H. habilis	1,900–600	South Africa	100 cm (3 ft 3 in)			3 individuals^[3]^{[lower-alpha 3]}	2010	2010
H. erectus	1,900–140^[4]^{[lower-alpha 4]}^[5]^{[lower-alpha 5]}	Africa, Eurasia	180 cm (5 ft 11 in)	60 kg (130 lb)	850 (early) – 1,100 (late)	Many^{[lower-alpha 6]}^{[lower-alpha 7]}	1891	1892
H. ergaster African H. erectus	1,800–1,300^[7]	East and Southern Africa			700–850	Many	1949	1975
H. antecessor	1,200–800	Western Europe	175 cm (5 ft 9 in)	90 kg (200 lb)	1,000	2 sites	1994	1997
H. heidelbergensis early H. neanderthalensis	600–300^{[lower-alpha 8]}	Europe, Africa	180 cm (5 ft 11 in)	90 kg (200 lb)	1,100–1,400	Many	1907	1908
H. cepranensis a single fossil, possibly H. heidelbergensis	c. 450^[8]	Italy			1,000	1 skull cap	1994	2003
H. longi	309–138^[9]	Northeast China			1,420^[10]	1 individual	1933	2021
H. rhodesiensis early H. sapiens	c. 300	Zambia			1,300	Single or very few	1921	1921
H. naledi	c. 300^[11]	South Africa	150 cm (4 ft 11 in)	45 kg (99 lb)	450	15 individuals	2013	2015
H. sapiens (anatomically modern humans)	c. 300–present^{[lower-alpha 9]}	Worldwide	150–190 cm (4 ft 11 in – 6 ft 3 in)	50–100 kg (110–220 lb)	950–1,800	(extant)	——	1758
H. neanderthalensis	240–40^[14]^{[lower-alpha 10]}	Europe, Western Asia	170 cm (5 ft 7 in)	55–70 kg (121–154 lb) (heavily built)	1,200–1,900	Many	1829	1864
H. floresiensis classification uncertain	190–50	Indonesia	100 cm (3 ft 3 in)	25 kg (55 lb)	400	7 individuals	2003	2004
Nesher Ramla Homo classification uncertain	140–120	Israel				several individuals	2021
H. tsaichangensis possibly H. erectus or Denisova	c. 100^{[lower-alpha 11]}	Taiwan				1 individual	2008(?)	2015
H. luzonensis	c. 67^[17]^[18]	Philippines				3 individuals	2007	2019
Denisova hominin	40	Siberia				2 sites	2000	2010^{[lower-alpha 12]}

Template:Homo

Template:Homo

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