The timeline of the evolutionary history of life represents the current scientific theory outlining the major events during the development of life on planet Earth. Dates in this article are consensus estimates based on scientific evidence, mainly fossils.

It has been suggested that portions of Geologic time scale (Table of geologic time) be split from it and merged into this article. (Discuss) (November 2023)

In biology, evolution is any change across successive generations in the heritable characteristics of biological populations. Evolutionary processes give rise to diversity at every level of biological organization, from kingdoms to species, and individual organisms and molecules, such as DNA and proteins. The similarities between all present day organisms imply a common ancestor from which all known species, living and extinct, have diverged. More than 99 percent of all species that ever lived (over five billion)^[1] are estimated to be extinct.^[2][3] Estimates on the number of Earth's current species range from 10 million to 14 million,^[4] with about 1.2 million or 14% documented, the rest not yet described.^[5] However, a 2016 report estimates an additional 1 trillion microbial species, with only 0.001% described.^[6]

There has been controversy between more traditional views of steadily increasing biodiversity, and a newer view of cycles of annihilation and diversification, so that certain past times, such as the Cambrian explosion, experienced maximums of diversity followed by sharp winnowing.^[7][8]

Species go extinct constantly as environments change, as organisms compete for environmental niches, and as genetic mutation leads to the rise of new species from older ones. At long irregular intervals, Earth's biosphere suffers a catastrophic die-off, a mass extinction,^[9] often comprising an accumulation of smaller extinction events over a relatively brief period.^[10]

The first known mass extinction was the Great Oxidation Event 2.4 billion years ago, which killed most of the planet's obligate anaerobes. Researchers have identified five other major extinction events in Earth's history, with estimated losses below:^[11]

• End Ordovician: 440 million years ago, 86% of all species lost, including graptolites
• Late Devonian: 375 million years ago, 75% of species lost, including most trilobites
• End Permian, The Great Dying: 251 million years ago, 96% of species lost, including tabulate corals, and most trees and synapsids
• End Triassic: 200 million years ago, 80% of species lost, including all conodonts
• End Cretaceous: 66 million years ago, 76% of species lost, including all ammonites, mosasaurs, plesiosaurs, pterosaurs, and nonavian dinosaurs

Smaller extinction events have occurred in the periods between, with some dividing geologic time periods and epochs. The Holocene extinction event is currently under way.^[12]

Factors in mass extinctions include continental drift, changes in atmospheric and marine chemistry, volcanism and other aspects of mountain formation, changes in glaciation, changes in sea level, and impact events.^[10]

In this timeline, Ma (for megaannum) means "million years ago," ka (for kiloannum) means "thousand years ago," and ya means "years ago."

Phanerozoic Eon

Main article: Phanerozoic

539 Ma – present

The Phanerozoic Eon (Greek: period of well-displayed life) marks the appearance in the fossil record of abundant, shell-forming and/or trace-making organisms. It is subdivided into three eras, the Paleozoic, Mesozoic and Cenozoic, with major mass extinctions at division points.

Palaeozoic Era

Main article: Paleozoic

This section needs additional citations for verification. (September 2022)

538.8 Ma – 251.9 Ma and contains the Cambrian, Ordovician, Silurian, Devonian, Carboniferous and Permian periods.

More information Date, Event ...

Date	Event
535 Ma	Major diversification of living things in the oceans: arthropods (e.g. trilobites, crustaceans), chordates, echinoderms, molluscs, brachiopods, foraminifers and radiolarians, etc.
530 Ma	The first known footprints on land date to 530 Ma.[74]
520 Ma	Earliest graptolites.[75]
511 Ma	Earliest crustaceans.[76]
505 Ma	Fossilization of the Burgess Shale
500 Ma	Jellyfish have existed since at least this time.
485 Ma	First vertebrates with true bones (jawless fishes).
450 Ma	First complete conodonts and echinoids appear.
440 Ma	First agnathan fishes: Heterostraci, Galeaspida, and Pituriaspida.
420 Ma	Earliest ray-finned fishes, trigonotarbid arachnids, and land scorpions.[77]
410 Ma	First signs of teeth in fish. Earliest Nautilida, lycophytes, and trimerophytes.
488–400 Ma	First cephalopods (nautiloids)[78] and chitons.[79]
395 Ma	First lichens, stoneworts. Earliest harvestmen, mites, hexapods (springtails) and ammonoids. The earliest known tracks on land named the Zachelmie trackways which are possibly related to icthyostegalians.[80]
375 Ma	Tiktaalik, a lobe-finned fish with some anatomical features similar to early tetrapods. It has been suggested to be a transitional species between fish and tetrapods.[81]
365 Ma	Acanthostega is one of the earliest vertebrates capable of walking.[82]
363 Ma	By the start of the Carboniferous Period, the Earth begins to resemble its present state. Insects roamed the land and would soon take to the skies; sharks swam the oceans as top predators,[83] and vegetation covered the land, with seed-bearing plants and forests soon to flourish. Four-limbed tetrapods gradually gain adaptations which will help them occupy a terrestrial life-habit.
360 Ma	First crabs and ferns. Land flora dominated by seed ferns. The Xinhang forest grows around this time.[84]
350 Ma	First large sharks, ratfishes, and hagfish; first crown tetrapods (with five digits and no fins and scales).
350 Ma	Diversification of amphibians.[85]
325-335 Ma	First Reptiliomorpha.[86]
330-320 Ma	First amniote vertebrates (Paleothyris).[87]
320 Ma	Synapsids (precursors to mammals) separate from sauropsids (reptiles) in late Carboniferous.[88]
305 Ma	The Carboniferous rainforest collapse occurs, causing a minor extinction event, as well as paving the way for amniotes to become dominant over amphibians and seed plants over ferns and lycophytes. First diapsid reptiles (e.g. Petrolacosaurus).
280 Ma	Earliest beetles, seed plants and conifers diversify while lepidodendrids and sphenopsids decrease. Terrestrial temnospondyl amphibians and pelycosaurs (e.g. Dimetrodon) diversify in species.
275 Ma	Therapsid synapsids separate from pelycosaur synapsids.
265 Ma	Gorgonopsians appear in the fossil record.[89]
251.9–251.4 Ma	The Permian–Triassic extinction event eliminates over 90-95% of marine species. Terrestrial organisms were not as seriously affected as the marine biota. This "clearing of the slate" may have led to an ensuing diversification, but life on land took 30 million years to completely recover.[90]

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Mesozoic Era

Main article: Mesozoic

This section needs additional citations for verification. (September 2022)

From 251.9 Ma to 66 Ma and containing the Triassic, Jurassic and Cretaceous periods.

More information Date, Event ...

Date	Event
250 Ma	Mesozoic marine revolution begins: increasingly well adapted and diverse predators stress sessile marine groups; the "balance of power" in the oceans shifts dramatically as some groups of prey adapt more rapidly and effectively than others.
250 Ma	Triadobatrachus massinoti is the earliest known frog.
248 Ma	Sturgeon and paddlefish (Acipenseridae) first appear.
245 Ma	Earliest ichthyosaurs
240 Ma	Increase in diversity of cynodonts and rhynchosaurs
225 Ma	Earliest dinosaurs (prosauropods), first cardiid bivalves, diversity in cycads, bennettitaleans, and conifers. First teleost fishes. First mammals (Adelobasileus).
220 Ma	Seed-producing Gymnosperm forests dominate the land; herbivores grow to huge sizes to accommodate the large guts necessary to digest the nutrient-poor plants.[citation needed] First flies and turtles (Odontochelys). First coelophysoid dinosaurs. First mammals from small-sized cynodonts, which transitioned towards a nocturnal, insectivorous, and endothermic lifestyle.
205 Ma	Massive Triassic/Jurassic extinction. It wipes out all pseudosuchians except crocodylomorphs, who transitioned to an aquatic habitat, while dinosaurs took over the land and pterosaurs filled the air.
200 Ma	First accepted evidence for viruses infecting eukaryotic cells (the group Geminiviridae).[91] However, viruses are still poorly understood and may have arisen before "life" itself, or may be a more recent phenomenon. Major extinctions in terrestrial vertebrates and large amphibians. Earliest examples of armoured dinosaurs.
195 Ma	First pterosaurs with specialized feeding (Dorygnathus). First sauropod dinosaurs. Diversification in small, ornithischian dinosaurs: heterodontosaurids, fabrosaurids, and scelidosaurids.
190 Ma	Pliosauroids appear in the fossil record. First lepidopteran insects (Archaeolepis), hermit crabs, modern starfish, irregular echinoids, corbulid bivalves, and tubulipore bryozoans. Extensive development of sponge reefs.
176 Ma	First Stegosaurian dinosaurs.
170 Ma	Earliest salamanders, newts, cryptoclidids, elasmosaurid plesiosaurs, and cladotherian mammals. Sauropod dinosaurs diversify.
168 Ma	First lizards.
165 Ma	First rays and glycymeridid bivalves. First vampire squids.[92]
163 Ma	Pterodactyloid pterosaurs first appear.[93]
161 Ma	Ceratopsian dinosaurs appear in the fossil record (Yinlong) and the oldest known eutherian mammal: Juramaia.
160 Ma	Multituberculate mammals (genus Rugosodon) appear in eastern China.
155 Ma	First blood-sucking insects (ceratopogonids), rudist bivalves, and cheilostome bryozoans. Archaeopteryx, a possible ancestor to the birds, appears in the fossil record, along with triconodontid and symmetrodont mammals. Diversity in stegosaurian and theropod dinosaurs.
131 Ma	First pine trees.
140 Ma	Orb-weaver spiders appear.
135 Ma	Rise of the angiosperms. Some of these flowering plants bear structures that attract insects and other animals to spread pollen; other angiosperms are pollinated by wind or water. This innovation causes a major burst of animal coevolution. First freshwater pelomedusid turtles. Earliest krill.
120 Ma	Oldest fossils of heterokonts, including both marine diatoms and silicoflagellates.
115 Ma	First monotreme mammals.
114 Ma	Earliest bees.[94]
112 Ma	Xiphactinus, a large predatory fish, appears in the fossil record.
110 Ma	First hesperornithes, toothed diving birds. Earliest limopsid, verticordiid, and thyasirid bivalves.
100 Ma	First ants.[95]
100–95 Ma	Spinosaurus, the largest theropod dinosaur, appears in the fossil record.[96]
95 Ma	First crocodilians evolve.[97]
90 Ma	Extinction of ichthyosaurs. Earliest snakes and nuculanid bivalves. Large diversification in angiosperms: magnoliids, rosids, hamamelidids, monocots, and ginger. Earliest examples of ticks. Probable origins of placental mammals (earliest undisputed fossil evidence is 66 Ma).
86–76 Ma	Diversification of therian mammals.[98][99]
70 Ma	Multituberculate mammals increase in diversity. First yoldiid bivalves. First possible ungulates (Protungulatum).
68–66 Ma	Tyrannosaurus, the largest terrestrial predator of western North America, appears in the fossil record. First species of Triceratops.[100]

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Cenozoic Era

Main article: Cenozoic

This section needs additional citations for verification. (September 2022)

66 Ma – present

More information Date, Event ...

Date	Event
66 Ma	The Cretaceous–Paleogene extinction event eradicates about half of all animal species, including mosasaurs, pterosaurs, plesiosaurs, ammonites, belemnites, rudist and inoceramid bivalves, most planktic foraminifers, and all of the dinosaurs excluding the birds.[101]
66 Ma-	Rapid dominance of conifers and ginkgos in high latitudes, along with mammals becoming the dominant species. First psammobiid bivalves. Earliest rodents. Rapid diversification in ants.
63 Ma	Evolution of the creodonts, an important group of meat-eating (carnivorous) mammals.
62 Ma	Evolution of the first penguins.
60 Ma	Diversification of large, flightless birds. Earliest true primates,[who?] along with the first semelid bivalves, edentate, carnivoran and lipotyphlan mammals, and owls. The ancestors of the carnivorous mammals (miacids) were alive.[citation needed]
59 Ma	Earliest sailfish appear.
56 Ma	Gastornis, a large flightless bird, appears in the fossil record.
55 Ma	Modern bird groups diversify (first song birds, parrots, loons, swifts, woodpeckers), first whale (Himalayacetus), earliest lagomorphs, armadillos, appearance of sirenian, proboscidean mammals in the fossil record. Flowering plants continue to diversify. The ancestor (according to theory) of the species in the genus Carcharodon, the early mako shark Isurus hastalis, is alive. Ungulates split into artiodactyla and perissodactyla, with some members of the former returning to the sea.
52 Ma	First bats appear (Onychonycteris).
50 Ma	Peak diversity of dinoflagellates and nannofossils, increase in diversity of anomalodesmatan and heteroconch bivalves, brontotheres, tapirs, rhinoceroses, and camels appear in the fossil record, diversification of primates.
40 Ma	Modern-type butterflies and moths appear. Extinction of Gastornis. Basilosaurus, one of the first of the giant whales, appeared in the fossil record.
38 Ma	Earliest bears.
37 Ma	First nimravid ("false saber-toothed cats") carnivores — these species are unrelated to modern-type felines. First alligators and ruminants.
35 Ma	Grasses diversify from among the monocot angiosperms; grasslands begin to expand. Slight increase in diversity of cold-tolerant ostracods and foraminifers, along with major extinctions of gastropods, reptiles, amphibians, and multituberculate mammals. Many modern mammal groups begin to appear: first glyptodonts, ground sloths, canids, peccaries, and the first eagles and hawks. Diversity in toothed and baleen whales.
33 Ma	Evolution of the thylacinid marsupials (Badjcinus).
30 Ma	First balanids and eucalypts, extinction of embrithopod and brontothere mammals, earliest pigs and cats.
28 Ma	Paraceratherium appears in the fossil record, the largest terrestrial mammal that ever lived. First pelicans.
25 Ma	Pelagornis sandersi appears in the fossil record, the largest flying bird that ever lived.
25 Ma	First deer.
24 Ma	First pinnipeds.
23 Ma	Earliest ostriches, trees representative of most major groups of oaks have appeared by now.[102]
20 Ma	First giraffes, hyenas, and giant anteaters, increase in bird diversity.
17 Ma	First birds of the genus Corvus (crows).
15 Ma	Genus Mammut appears in the fossil record, first bovids and kangaroos, diversity in Australian megafauna.
10 Ma	Grasslands and savannas are established, diversity in insects, especially ants and termites, horses increase in body size and develop high-crowned teeth, major diversification in grassland mammals and snakes.
9.5 Ma [dubious – discuss]	Great American Interchange, where various land and freshwater faunas migrated between North and South America. Armadillos, opossums, hummingbirds Phorusrhacids, Ground Sloths, Glyptodonts, and Meridiungulates traveled to North America, while horses, tapirs, saber-toothed cats, jaguars, bears, coaties, ferrets, otters, skunks and deer entered South America.
9 Ma	First platypuses.
6.5 Ma	First hominins (Sahelanthropus).
6 Ma	Australopithecines diversify (Orrorin, Ardipithecus).
5 Ma	First tree sloths and hippopotami, diversification of grazing herbivores like zebras and elephants, large carnivorous mammals like lions and the genus Canis, burrowing rodents, kangaroos, birds, and small carnivores, vultures increase in size, decrease in the number of perissodactyl mammals. Extinction of nimravid carnivores. First leopard seals.
4.8 Ma	Mammoths appear in the fossil record.
4.5 Ma	Marine iguanas diverge from land iguanas.
4 Ma	Australopithecus evolves. Stupendemys appears in the fossil record as the largest freshwater turtle, first modern elephants, giraffes, zebras, lions, rhinoceros and gazelles appear in the fossil record
3.6 Ma	Blue whales grow to modern size.
3 Ma	Earliest swordfish.
2.7 Ma	Paranthropus evolves.
2.5 Ma	Earliest species of Arctodus and Smilodon evolve.
2 Ma	First members of genus Homo, Homo Habilis, appear in the fossil record. Diversification of conifers in high latitudes. The eventual ancestor of cattle, aurochs (Bos primigenus), evolves in India.
1.7 Ma	Australopithecines go extinct.
1.2 Ma	Evolution of Homo antecessor. The last members of Paranthropus die out.
1 Ma	First coyotes.
810 ka	First wolves
600 ka	Evolution of Homo heidelbergensis.
400 ka	First polar bears.
350 ka	Evolution of Neanderthals.
300 ka	Gigantopithecus, a giant relative of the orangutan from Asia dies out.
250 ka	Anatomically modern humans appear in Africa.[103][104][105] Around 50 ka they start colonising the other continents, replacing Neanderthals in Europe and other hominins in Asia.
70 ka	Genetic bottleneck in humans (Toba catastrophe theory).
40 ka	Last giant monitor lizards (Varanus priscus) die out.
35-25 ka	Extinction of Neanderthals. Domestication of dogs.
15 ka	Last woolly rhinoceros (Coelodonta antiquitatis) are believed to have gone extinct.
11 ka	Short-faced bears vanish from North America, with the last giant ground sloths dying out. All Equidae become extinct in North America. Domestication of various ungulates.
10 ka	Holocene epoch starts[106] after the Last Glacial Maximum. Last mainland species of woolly mammoth (Mammuthus primigenus) die out, as does the last Smilodon species.
8 ka	The Giant Lemur dies out.

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