Urocyclidae

Urocyclidae

Urocyclidae

Family of gastropods

Urocyclidae is a family of air-breathing land snails, semi-slugs and land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Helicarionoidea, (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).[2]

Quick Facts Scientific classification ...

Some species within this family make and use love darts before mating.[citation needed]

Distribution

The Urocyclidae are distributed in the Afrotropical realm, including Madagascar, the Comores and the islands of the Gulf of Guinea.[3][4]

Taxonomy

The family Urocyclidae consists of 3 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005),[2] that follows classification of Urocyclinae by Van Goethem (1977):[5]

  • subfamily Rhysotininae Schileyko, 2002
  • subfamily Sheldoniinae Connolly, 1925 (1912)[6] - synonyms: Peltatinae Godwin-Austen, 1912;[7] Trochonanininae Connolly, 1912;[8] Trochozonitinae Iredale, 1914;[9] Ledoulxiinae Pilsbry, 1919;[10] Gymnarionidae Von Mol, 1970;[11] Rhysotinidae Schileyko, 2002; Zonitarionini Schileyko, 2002; Acantharionini Schileyko, 2002
  • subfamily Urocyclinae Simroth, 1889
    • tribe Urocyclini Simroth, 1889 - synonyms: Atoxonini Schileyko, 2002; Buettneriini Schileyko, 2002
    • tribe Dendrolimacini Van Goethem, 1977[5]
    • tribe Leptichnini Van Goethem, 1977[5]
    • tribe Upembellini Van Goethem, 1977[5]

Genera

Genera within the family Urocyclidae include:

unsorted
  • † Koruella Pickford, 2009
Subfamily Rhysotininae Schileyko, 2002
  • Rhysotina Ancey, 1887[2]
  • Thomeonanina Germain, 1909 : synonym of Rhysotina Ancey, 1887 (objective synonym)
Subfamily Sheldoniinae
  • Acantharion Binder & Tillier, 1985[2]
  • Africarion Godwin-Austen, 1883
  • Amatarion Van Mol, 1970
  • Angustivestis Pilsbry, 1919
  • Aspidelus Morelet, 1884
  • Aspidotomium Degner, 1932
  • Belonarion Pilsbry, 1919
  • Bloyetia Bourguignat, 1890
  • Camerunarion Van Mol, 1970
  • Carinazingis van Bruggen & de Winter, 1990
  • Centrafricarion Van Mol, 1970 [11]
  • Chlamydarion Van Mol, 1968
  • Colparion Laidlaw, 1938
  • Degneria Verdcourt, 1956
  • Entagaricus Pilsbry, 1919
  • Estria Poirier, 1887
  • Falloonella Preston, 1914
  • Fuellebornia Verdcourt, 1998
  • Granularion Germain, 1912
  • Gymnarion Pilsbry, 1919[2]
  • Hamya Bourguignat, 1885 †
  • Kerkophorus Godwin-Austen, 1912
  • Lacrimarion Connolly, 1939
  • Malagarion Tillier, 1979
  • Mesafricarion Pilsbry, 1919
  • Microkerkus Godwin-Austen, 1912
  • Montanobloyetia Verdcourt, 1961
  • Morrumbalia Verdcourt, 1998
  • Paragranularion Van Mol, 1970
  • Percivalia Preston, 1914
  • Plicatonanina Verdcourt, 1961
  • Principicochlea D. Holyoak & G. Holyoak, 2020
  • Principitrochoidea D. Holyoak & G. Holyoak, 2020
  • Ptilototheca Herbert, 2016
  • Rhopalogonium Degner, 1932
  • Selatodryas Herbert, 2017
  • Senegalarion Van Mol, 1970
  • Sheldonia Ancey, 1887 - the type genus of the subfamily Sheldoniinae[2][12]
  • Sjostedtina Verdcourt, 1961
  • Sylvarion Van Mol, 1970
  • Thapsia Albers, 1860
  • Thielarion Van Mol, 1970
  • Thomithapsia D. Holyoak & G. Holyoak, 2020
  • Thomitrochoidea D. Holyoak & G. Holyoak, 2020
  • Tresia Van Goethem, 1975
  • Trochonanina Mousson, 1869[2]
  • Trochozonites Pfeffer, 1883[2]
  • Tropidocochlion Verdcourt, 1998
  • Verdcourtia Van Mol, 1970
  • Verrucarion Van Mol, 1970 [13]
  • Zingis E. von Martens, 1878
  • Zonitarion Pfeffer, 1883[2]
  • Ledoulxia Bourguignat, 1885:[2] synonym of Trochonanina Mousson, 1869
  • Peltatus Godwin-Austen, 1908:[2] synonym of Sheldonia Ancey, 1887
Subfamily Urocyclinae

Tribe Dendrolimacini

  • Dendrolimax Heynemann, 1868 - the type genus of the tribe Dendrolimacini[2]

Tribe Leptichnini

  • Leptichnus Simroth, 1896 - the type genus of the tribe Leptichnini[2]

Tribe Upembellini

  • Leptichnoides Van Goethem, 1975
  • Upembella Van Goethem, 1969 - the type genus of the tribe Upembellini[2]

tribe Urocyclini

  • Anisotoxon Van Goethem, 1975
  • Atoxon Simroth, 1888[2]
  • Atoxonoides Van Goethem, 1973
  • Atrichotoxon Simroth, 1910
  • Buettneria Simroth, 1910[14]
  • Bukobia Simroth, 1896
  • Elisolimax Cockerell, 1893
  • Emphysetes Verdcourt, 2003
  • Microcyclus Simroth, 1896
  • Nupnus Van Goethem, 1975
  • Pembatoxon Van Goethem, 1975
  • Phaneroporus Simroth, 1888
  • Polytoxon Simroth, 1897
  • Pseudatoxon Van Goethem, 1975
  • Tanzalimax Rowson, Paustian & Van Goethem, 2017
  • Trichotoxon Simroth, 1888
  • Udzungwalimax Rowson, Paustian & Van Goethem, 2017
  • Urocyclus Gray, 1864 - the type genus of the family Urocyclidae[2]

Cladogram

The following cladogram shows the phylogenic relationship of this family to other families in the limacoid clade:[3]

 limacoid clade 

References

  1. [1]
    Simroth H. (1899). "Beiträge zur Kenntniss der Nacktschnecken". Nova Acta, Kaiserlich Leopoldinish-Carolinische Deutsche Akademie der Naturforscher (= Nova Acta Academiae Caesareae Leopoldino-Carolinae Germanicae Naturae Curiosorum) 54(1): 1-91, plates 1-4. p. 62.
  2. [2]
    Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. 47 (1–2). Hackenheim, Germany: ConchBooks: 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
  3. [3]
    Hausdorf B. (2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". Journal of Biogeography 27(2): 379-390. doi:10.1046/j.1365-2699.2000.00403.x, JSTOR.
  4. [4]
    Herbert, Dai; Kilburn, Dick (2004). Field guide to the land snails and slugs of eastern south Africa (1st ed.). Pietermaritzburg :Natal Museum: Natal Museum. ISBN 0-620-32415-5.
  5. [5]
    Van Goethem J. (1977). "Révision systématique des Urocyclinae (Mollusca, Pulmonata, Urocyclidae)". Musée Royal de l'Afrique Centrale , Annales, Sciences zoologiques 218: 355 pp., 4 plates.
  6. [6]
    Connolly (1925). Annals and Magazine of Natural History (9)15: 467.
  7. [7]
    Godwin-Austen H. H. (1912). "A review of South-African Land-Mollusca belonging to the Family Zonitidae". Annals and Magazine of Natural History (8)9: 122-139. Plate 1-7. p. 124.
  8. [8]
    Connolly (1912). Annals of the South African Museum 11(3): 101.
  9. [9]
    Iredale (1914). Proceedings of the Malacological Society of London 11(2): 122.
  10. [10]
    Pilsbry H. A. (1919). Bulletin of the American Museum of Natural History 40: 245.
  11. [11]
    Van Mol J. J. (1970). "Révision des Urocyclidae (Mollusca, Gastropoda, Pulmonata). Anatomie - systématique - zoogéographie". Annales Musée Royal de l’Afrique Centrale, Sciences Zoologiques 180: 1-231.
  12. [12]
    Herbert D. G. (2004). Sheldonia puzeyi. 2006 IUCN Red List of Threatened Species. Downloaded on 7 August 2007.
  13. [13]
    de Winter A. J. (1987). "Verrucarion demeryi, a new semi-slug from Sierra Leone (Mollusca: Pulmonata: Urocyclidae)". Basteria 51(4-6): 87-93. abstract.
  14. [14]
    Simroth H. (1910). In: Voeltzkow Reise O.-Afr., Wiss. Ergebn. 2(5): 611.
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